Lasso, Carlos A., José Iván Mojica, José Saulo Usma, Javier A. Maldonado-Ocampo, Carlos DoNascimiento, Donald C. Taphorn, Francisco Provenzano, et al. 2004. “Fish Species of the Orinoco Basin. Part I: Species List and Distribution According to Subbasins”. Biota Colombiana 5 (2). https://revistas.humboldt.org.co/index.php/biota/article/view/141.